Publications

Publications
2010-2012

2011-12

1. Cable, J. Ogle, K., Welker, J.M. and 10 additional authors. (2011) The temperature response of soil respiration: A 7 desert synthesis. Biogeochemistry. 103: 71-90.

2. Cahoon, S., Sullivan, P.F., Post, E., and Welker, J.M. Herbivores constrain Arctic CO2 uptake and surpress C cycling responses to warming. Global Change Biology. (In Press)

3. Chimner, R., and Welker, J.M. Influence of grazing and precipitation on ecosystem carbon cycling in a mixedgrass prairie. Pastorial Science. (In Press)

4. Ives, S., Sullivan, P.F., Dail, R., Berg, E. and Welker, J.M. Seasonal and inter-annual patterns in the magnitudes of CO2 exchange along a hydrologic gradient in the Kenai Lowlands, AK: Feedback implications of wetland drying and vegetation succession. Ecohydrology. (In Press)

5. Rogers, M., Sullivan, P. and Welker, J.M. (2011) Experimental increases in snow depth delays the seasonality but enhances the magnitudes of vegetation traits and trace gas exchanges in the High Arctic of NW Greenland. Arctic, Antarctic and Alpine Research. 43: 95-106.

6. Tape, K. D., Verbyla, D. and Welker, J.M. Erosion decline in Arctic Alaska uplands since 1980: the influence of shrubs, runoff, and permafrost . JGR Biogeosciences. (In Press)

7. Werner, C., Welker, J.M., et al. (2011) Linking carbon and water cycles using stable isotopes across scales: Progress and challenges. Biogeosciences Discussions. 8: 2659-2719. (On-line Journal)

8. Wilgenberg, S. L., Hobson, K. A., Brewster, K, and Welker, J.M. Addressing uncertainty in assessing dispersal in threatened migratory species using stable hydrogen (δD) analysis of feathers. Conservation Biology. (In Press)

9. Miles, G.R., Mitchell, M.J., Mayer, B., Likens, G., and Welker, J. Long-term analysis of Hubbard Brook stable oxygen isotope ratios of streamwater and precipitation sulfate. Biogeochemistry. doi:10.1007/s10533-011-9670-3. (In Press)

10. Myers-Smith, I.H., Forbes, B.C., Wilmking, M., Hallinger, M., Lantz, T., Blok, D., Tape, K.D.*, Macias-Fauria, M., Sass-Klassen-U., Lévesque, E., Boudreau, S., Ropars, P., Hermanutz, L., Trant, A., Siegwart Collier, L., Weijers, S., Rozema, J., Rayback, S.A., Schmidt, N.M., Schaepman-Strub, G., Wipf, S., Rixen, C., Ménard, C.B., Venn, S., Goetz, S., Andreu-Hayles, L., Elmendorf, S., Ravolainen, V., Welker, J., Grogan, P., Epstein, H.E., and Hik, D.S. (2011) Shrub expansion in tundra ecosystems: Dynamics, impacts and research priorities. Environmental Research Letters. 6(4):045509. doi:10.1088/1748-9326/6/4/045509

2010

9. Chimner, R., Welker, J.M., Morgan, J., LeCain, D. and Reader, J. (2010) Experimental manipulations of winter snow cover and summer rain influence ecoystem C cycling in a Mixedgrass Prairie. Ecohydrology. 3: 284-297.

10. Czimczik, C., and Welker, J.M. (2010) Age and sources of soil respiration from High Arctic tundra in NW Greenland. Arctic, Antarctic and Alpine Research. 42: 342-350.

11. Liu, Z., Bowen, G., and Welker, J.M. (2010) Precipitation isotope (δ18O) gradients depict modern atmospheric circulation over the conterminous United States. Journal of Geophysical Research. 115: D22120, doi:10.1029/2010JD014175.

12. Nowinski, N., Taneva, L., Trumbore, S., and Welker, J.M. (2010) Decomposition of old organic matter as a result of a deeper active layer in a snow manipulation experiment. Oecologia 163: 785-792.

13. Stevenson, B.A., Kelly, E., McDonald, E., Busacca. A., and Welker, J.M. (2010) Oxygen isotope ratios in Holocene carbonates across a climatic gradient eastern Washington State, USA: Evidence for seasonal effects on pedogenic mineral isotopic composition. The Holocene. 20: 575-583.


14. Sullivan, P.F., Arens, S., Sveinbjörnsson, B. and Welker, J.M. (2010) Modeling the seasonality of belowground respiration along an elevational gradient in the Chugach Mountains, south central Alaska. Biogeochemistry. 101: 61-75.

15. Vachon, R.W., Welker, J.M., White, J.W.C., and Vaughn, B.H. (2010) Monthly precipitation isoscapes (δ18O) of the United States: Connections with surface temperatures, moisture source conditions, and air mass trajectories. Journal of Geophysical Research. 115: D21126. doi:10.1029/2010JD014105.

16. Vachon, R.W., Welker, J.M., White, J.W., and Vaughn, B.H. (2010) Moisture source temperatures and precipitation δ18O-temperature relationships across the US. Water Resources Research. 46: W07523.

Submitted Papers

17. Kikuchi, C., Ferre, T.P., and Welker, J.M. Groundwater-surface water interactions in an Alaskan freshwater stream. Journal of Hydrology. (Submitted 2011)

18. Miles, G.R., Mitchell, M.J., Mayer, B., Likens, G., and Welker, J.M. Long-term analysis of Hubbard Brook stable oxygen isotope ratios of stream water and precipitation sulfate. Biogeochemistry. (Submitted 2011)

19. Tape, K.D., Hallinger, M., Reuss, R. and Welker, J.M. Landscape heterogeneity of shrub expansion in Arctic Alaska: Patterns and processes controlling vegetation change. Global Change Biology. (Submitted 2011)

20. Springer, G.S., Rowe, H.D., Hardt, B., Edwards, R.L., Welker, J.M., and Cheng, H. A tropical origin for seasonality and the abrupt Mid to Late Holocene climatic transition in eastern North America based on a 7000 year δ18O record in stalagmite calcite. Holocene. (Submitted 2010)

21. von Biela, V.R., Zimmerman, C.E., Cohn, B.R. , and Welker, J.M. Evidence of multiple carbon sources and prey communities in young-of-the-year Arctic cisco (Coregonus autumnalis) using δ13C, δ15N and diet analyses. Polar Biology. (Submitted 2011)

All Publications organized by Themes (108)

Arctic Ecosystem & Physiological Ecology

1. Rogers, M., Sullivan, P. and Welker, J.M. (2011) Experimental increases in snow depth delays the seasonality but enhances the magnitudes of vegetation traits and trace gas exchanges in the High Arctic of NW Greenland. Arctic, Antarctic and Alpine Research. 43: 95-106.

2. Tape, K. D., Hallinger, M., Reuss, R. and Welker, J.M. Landscape heterogeneity of shrub expansion in Arctic Alaska: Patterns and processes controlling vegetation change. Global Change Biology. (Submitted 2011)

3. Tape, K. D., Verbyla, D. and Welker, J.M. Erosion decline in Arctic Alaska uplands since 1980: The influence of shrubs, runoff, and permafrost. Biogeosciences. (In Press)

4. Nowinski, N, Taneva, L., Trumbore, S., and Welker, J.M. (2010) Decomposition of old organic matter as a result of a deeper active layer in a snow manipulation experiment. Oecologia. 163: 785-792.

5. Czimczik, C., and Welker, J.M. (2010) Age and sources of soil respiration from High Arctic tundra in NW Greenland. Arctic, Antarctic and Alpine Research. 42: 342-350.

6. Sullivan, P.F., Arens, S., Sveinbjörnsson, B. and Welker, J.M. (2010) Modeling the seasonality of belowground respiration along an elevational gradient in the Chugach Mountains, south central Alaska. Biogeochemistry. 101: 61-75.

7. Taneva, L., Sullivan, P.F., and Welker, J.M. Identifying the cause and biogeochemical consequences of high shrub abundance in low arctic tundra. Global Climate Change. (Submitted 2010)

8. Post, E., Forchhammer, M.C., Welker, J.M., et al. (2009) Ecological dynamics across the Arctic associated with recent climate change. Science. 325: 1355-1358.

9. Sullivan P.F., Welker, J.M., Arens, S.J.T., and Sveinbjörnsson, B. (2008) Continuous estimates of CO2 efflux from Arctic and boreal soils during the snow-covered season in Alaska. JGR Biogeosciences. 113: G04009.

10. Walker, D.A., Epstein, H.E., and Welker, J.M. (2008) Introduction to the special section: Biocomplexity in Arctic terrestrial environments. Journal of Geophysical Research. G03S14.

11. Sullivan, P.F, Welker, J.M., Steltzer, H., Sletten, R., Hagedorn, B., Arens, S., and Horwath, J.L. (2008) Energy and water additions give rise to simple responses in plant canopy and soil microclimates of a high arctic ecosystem. Journal of Geophysical Research. G03S08.

12. Arens, S., Sullivan, P.F., and Welker, J.M. (2008) Nonlinear responses to nitrogen and strong interactions with nitrogen and phosphorus additions drastically alter the structure and function of a High Arctic ecosystem. Journal of Geophysical Research. G03S09.

13. Steltzer H., Hufbauer, R.A, Welker, J.M., Casalis, M., Sullivan, P.F., and Chimner, R.A. (2008) Frequent sexual reproduction and high intraspecific variation in Salix arctica: Implications for a terrestrial feedback to climate change in the High Arctic. Journal of Geophysical Research. G03S10.

14. Sullivan, P.F., Arens, S.J.T., Chimner, R.A., and Welker, J.M. (2008) Temperature and microtopography interact to control carbon cycling in a High Arctic fen. Ecosystems. 11: 61-76.

15. Oberbauer, S., Tweedie, C., Welker, J.M., et al. (2007) Tundra CO2 fluxes in response to experimental warming across latitudinal and moisture gradients. Ecological Monographs. 77: 221-238.

16. Sullivan, P.F., and Welker, J.M. (2007) Variation in leaf physiology of Salix arctica within and across ecosystems in the High Arctic: Test of a dual isotope conceptual model. Oecologia. 151: 372-386.

17. Cornelissen, J., Bodegom, J., Aerts, R., Callaghan, T., Welker, J.M., et al. (2007) Global negative vegetation feedback to climate warming responses to leaf litter decomposition rates in cold biomes. Ecology Letters. 10: 619-627.

18. Sullivan, P.F., Sommerkorn, M., Rueth, H., Nadelhoffer, K., Shaver, G. and Welker, J.M. (2007) Climate and species affect fine root production with long-term fertilization in acidic tussock tundra near Toolik Lake, Alaska. Oecologia. 153: 643-652.

19. Steltzer, H., and Welker, J.M. (2006) Modeling the effect of photosynthetic vegetation properties on the NDVI-LAI relationship. Ecology. 87: 2765-2772.

20. Schimel, J., Fahnestock, J., Michaelson, G., Milkan, C., Ping, C, Romanovsky, V and Welker, J.M. (2006) Cold-season production of CO2 in Arctic soils: Can laboratory and field estimates be reconciled through a simple modeling approach. Arctic, Antarctic and Alpine Research. 33: 120-125.

21. Sturm, M., Schimel, J., Michaelson, G., Welker, J.M., Oberbauer, S. F., Liston, G., Fahnestock, J. T., and Romanovsky, V. E. (2005) Winter biological processes could help convert Arctic tundra to shrubland. Bioscience. 55, 17-26.

22. Chapin, F.S., Strum, M., Serreze, M., McFadden, J., Key, J., Lloyd, A., McGuire, D, Rupp T., Lynch, A., Schimel, J., Beringer, J., Epstein, H., Hinzman, L., Jia, G., Ping, C-L., Tape, K., Chapman, W., Euskirchen, E., Thompson, C., Welker, J.M., and Walker, D. (2005) Amplification of Arctic summer warming by terrestrial ecosystem changes. Science. 310: 657-660.

23. Hinzman, L.D., Oechel, W.C., Sturm, M., Welker, J.M., Nelson, F.E., Romanovsky, V.E., Chapin, F.S., et al. (2005) Evidence and implications of recent climate change in northern Alaska and other Arctic regions. Climatic Change. 72(3): 251-298. doi:10.1007/s10584-005-5352-2.

24. Sullivan, P.F. and Welker, J.M. (2005) Warming chambers stimulate early season growth of an Arctic sedge: Results of a minirhizotron field study. Oecologia. 142: 616-626.

25. Welker, J.M., Fahnestock, J.T., Sullivan, P. and Chimner, R.A. (2005) Leaf mineral nutrition of Arctic plants in response to long-term warming and deeper snow in northern Alaska. Oikos. 109: 167-177.

26. Welker, J.M., Rayback, S., and Henry, G. (2005) Arctic and North Atlantic Oscillation phase changes are recorded in the isotopes (δ18O & δ13C) of Cassiope tetragona plants. Global Change Biology. 11: 1-6.

27. Welker, J.M., Fahnestock, J.T., Henry, G., O’dea, K., and Chimner, R. (2004) CO2 exchange in the Canadian High Arctic: Response to long-term warming. Global Change Biology. 10: 1981-(1995)

28. Schimel, J.S., Bilbrough, C.B., and Welker, J.M. (2004) The effect of changing snow cover on year-round soil nitrogen dynamics in Arctic tundra ecosystems. Soil Biology and Biochemistry. 36: 217-227.

29. Welker, J.M., Jonsdottir, I., and Fahnestock, J.T. (2003) Isotopic (δ13C and δ15N) characteristics of Carex plants and populations along the Eurasian Coastal Arctic. Polar Biology. 27: 29-37.

30. Welker, J.M., Fahnestock, J.T., Henry, G., Odea, K. and Piper, R. (2002) Microbial activity discovered in previously ice-entombed Arctic ecosystems. EOS Transactions American Geophysical Union. 83: 281 & 284.

31. Welker, J.M., Fahnestock, J.T., and Jones, M.H. (2000) Annual CO2 flux from dry and moist Arctic tundra: Field responses to increases in summer temperature and winter snow depth. Climatic Change. 44: 139-150.

32. Fahnestock, J.T., Povirk, K.A. and Welker, J.M. (2000) Abiotic and biotic effects of increased litter accumulation in arctic tundra. Ecography. 23: 623-631.

33. Bilbrough, C.J., Welker, J.M., and Bowman, W.D (2000) Early-spring N uptake by snow covered plants: A comparison of Arctic and alpine plant function under snowpack. Arctic, Antarctic and Alpine Research. 32: 404-411.

34. Jones, M.H., Fahnestock, J.T., Stahl, P.D., and Welker, J.M. (2000) A note on summer CO2 flux, soil organic matter and soil microbial biomass from different high Arctic ecosystem types in northwestern Greenland. Arctic, Antarctic and Alpine Research. 32: 104-106.

35. Fahnestock, J.T., Jones, M.H., Brooks, P.D., and Welker, J.M. (1999) Significant CO2 emissions from tundra soils during winter: Implications for annual carbon budgets of arctic communities. Global Biogeochemical Cycles. 13: 775-779.

36. Jones, M.H., Fahnestock, J.T. and Welker, J.M. (1999) Early and late winter CO2 efflux from Arctic tundra in the Kuparuk River watershed, Alaska. Arctic, Antarctic and Alpine Research. 31: 187-190.

37. Arft, A.M., Walker, M.D., Gurevitch, J., Welker J.M., et al. (1999) Response patterns of tundra plant species to experimental warming: A meta-analysis of the International Tundra Experiment. Ecological Monographs. 69: 491-511.

38. Welker, J.M., Brown, K.B., and Fahnestock, J.T. (1999) CO2 flux in Arctic and alpine dry tundra: Comparative field responses under ambient and experimentally warmed conditions. Arctic, Antarctic and Alpine Research. 31: 308-313.

39. Walker, M.D., Walker, D.A., Welker, J.M., Arft, A.M., Bardsley, T., Brooks, P.D., Fahnestock, J.T., Jones, M.H., Losleben, M., Parsons, A.N., Seastedt, T.R. and Turner, P.L. (1999) Long-term experimental manipulation of winter snow regime and summer temperature in Arctic and alpine tundra. Hydrological Processes. 13: 2315-2330.

40. Jones, M.H., Fahnestock, J.T., Walker, D.A., Walker, M.D., and Welker, J.M. (1998) Carbon dioxide fluxes in moist and dry Arctic tundra during the snow-free season: Responses to increases in summer temperature and winter snow accumulation. Arctic and Alpine Research. 30: 373-380.

41. Fahnestock, J.T., Jones, M.H., Brooks, P.D., Walker, D.A., and Welker, J.M. (1998) Winter and early spring CO2 flux from tundra communities of northern Alaska. Journal of Geophysical Research. 102(D22): 29925-29931.

42. Welker, J.M., Molau, U., Parsons, A.N., Robinson, C. and Wookey, P.A. (1997) Response of Dryas octopetala to ITEX manipulations: A synthesis with circumpolar comparisons. Global Change Biology. 3(suppl. 1): 61-73.

43. Walsh, N., McCabe, T., Welker, J.M., and Parsons, A.N. (1997) Experimental manipulations of snow-depth: Effects on nutrition content of caribou forage. Global Change Biology. 3(suppl. 1): 158-164.

44. Robinson, C.H., Wookey, P.A., Parson, A.N., Welker, J.M., Callaghan, T.V., Press, M.C., and Lee, J.A. (1995) The response of plant litter decomposition, soil nutrient release and nitrogen mineralization to simulate climate change in a high Arctic polar semi-desert and a subarctic dwarf shrub heath. Oikos. 74: 503-512.

45. Wookey, P.A., Robinson, C.H., Parsons, A.N., Welker, J.M., Press, M. Callaghan, T.V., and Lee, J. A. (1995) Environmental constraints on the growth and performance of Dryas octopetala ssp. at a high Arctic polar semi-desert. Oecologia. 102: 478-489.

46. Parsons, A.N., Wookey, P.A., Welker, J.M., Press, M.C., Callaghan T.V., and Lee, J.A. (1995) Growth of Calamagrostis lapponica in response to simulated environmental change in the Subarctic. Oikos. 72: 61-66.

47. Rieley, G., Welker, J.M., Eglinton, G., and Callaghan, T.V. (1995) Epicuticular waxes of Arctic plants: Compositional differences in relation to winter snow cover. Phytochemistry. 38: 45-52.

48. Parsons, A.N., Welker, J.M., Wookey, P.A., Press, M.C. Callaghan, T.V., and Lee, J.A. (1994) Growth responses of four dwarf shrubs to simulated climate change. Journal of Ecology. 82: 307-318.

49. Wookey, P.A., Welker, J.M., Parsons, A.N., Press, M.C., Callaghan, T.V., and Lee, J.A. (1994) Differential growth allocation and photosynthetic responses of Polygonum viviparum L. to simulated environmental change at a high Arctic polar semi-desert. Oikos. 70: 131-139.

50. Wookey, P.A., Parsons, A.N., Welker, J.M., Potter, J.C., Callaghan, T.V., Lee, J.A., and Press, M.C. (1993) Comparative responses of subarctic and high Arctic ecosystems to simulated climate change. Oikos. 67:490-502.

51. Welker, J.M., Wookey, P., Parsons, A. P., Callaghan, T. V., Press, M. C., and Lee, J. A. (1993) Leaf carbon isotope discrimination and demographic responses of Dryas octopetala to water and temperature manipulations in a high Arctic polar semi-desert, Svalbard. Oecologia. 95:463-470.

Isotope Biogeochemistry

52. Miles, G.R., Mitchell, M.J., Mayer, B., Likens, G., and Welker, J.M. Long-term analysis of Hubbard Brook stable oxygen isotope ratios of stream water and precipitation sulfate. Biogeochemistry. (Resubmitted 2011)

53. Kikuchi, C., Ferre, T.P., and Welker, J.M. Groundwater-surface water interactions in an Alaskan freshwater stream. Journal of Hydrology. (Submitted 2011)

54. Werner, C., Welker, J.M., et al. (2011) Linking carbon and water cycles using stable isotopes across scales: Progress and challenges. Biogeosciences Discussions. 8: 2659-2719. (On-line Journal)

55. von Biela, V.R., Zimmerman, C.E. , Cohn, B.R. , and Welker, J.M. Evidence of multiple carbon sources and prey communities in young-of-the-year Arctic cisco (Coregonus autumnalis) using δ13C, δ15N and diet analyses. Polar Biology. (Submitted 2011)

56. Wilgenberg, S.L., Hobson, K.A., Brewster, K., and Welker, J.M. Addressing uncertainty in assessing dispersal in threatened migratory species using stable hydrogen (δD) analysis of feathers. Conservation Biology. (Submitted 2011)

57. Stevenson, B.A., Kelly, E., McDonald, E., Busacca, A., and Welker, J.M. (2010) Oxygen isotope ratios in Holocene carbonates across a climatic gradient eastern Washington State, USA: Evidence for seasonal effects on pedogenic mineral isotopic composition. The Holocene. 20: 575-583.


58. Vachon, R.W., Welker, J.M. White, J.W.C., and Vaughn, B.H. (2010) Monthly precipitation isoscapes (δ18O) of the United States: Connections with surface temperatures, moisture source conditions, and air mass trajectories. Journal of Geophysical Research. 115: D21126. doi:10.1029/2010JD014105.

59. Vachon, R.W., Welker, J.M., White, J.W., and Vaughn, B.H. (2010) Moisture source temperatures and precipitation δ18O-temperature relationships across the US. Water Resources Research. 46: W07523

60. Liu, Z., Bowen, G., and Welker, J.M. (2010) Precipitation isotope (δ18O) gradients depict modern atmospheric circulation over the conterminous United States. Journal of Geophysical Research. 115: D22120. doi:10.1029/2010JD014175.

61. Springer, G.S., Rowe, H.D., Hardt, B., Edwards, R.L., Welker, J.M., and Cheng, H. (2010) A tropical origin for seasonality and the abrupt Mid to Late Holocene climatic transition in eastern North America based on a 7000 year δ18O record in stalagmite calcite. Holocene. (Submitted)

62. Vachon, R.W., and Welker, J.M. New insights into the isotope thermometer gleaned from a dissection of the Rayleigh Distillation Model (Under Peer Review)

63. Craine, J.M., Elmore, A.J., Aidar, M.P.M., Amundson, R., Bustamante, M., Coetsee, C., Dawson, T., Hawkins, H., Hobbie, E., Kahmen, A., Koba, K., Mack, M., Makarov, M., McLauchlan, K., Michelsen, A., Nardoto, G., Pardo, L., Peñuelas, J., Reich, P., Schuur, E., Stock, W., Tateno, R., Templer, P., Virginia, R., Welker, J.M., and Wright, I. (2009) Global patterns of foliar nitrogen isotopes and their relationships with climate, mycorrhizal fungi, foliar nutrient concentrations, and nitrogen availability. New Phytologist. 183: 980-992.

64. Sjostrom, D., and Welker, J.M. (2009) Storm-track trajectories control the isotope geochemistry of precipitation in the Eastern US. Exploration Geology-Isobalance Special Issue. 102: 103-112

65. Still, C.J., Riley, W.J., Biraud, S.C., Noone, D.C., Buenning, N.H., Randerson J.T., Torn, M.S., Welker, J.M., White, J.W.C., Vachon, R., Farquhar, G.D., and Berry, J.A. (2009) The influence of clouds and diffuse radiation on ecosystem-atmosphere CO2 and CO18O exchanges. Biogeosciences. 7: 210-217.

66. Vachon, R., White, J.W., Welker, J.M., and Gutman, S. (2007) Seasonality of precipitation affects the annual isotopic (δ18O) values across the United States: A sensitivity study. Geophysical Research Letters. 34: L21707.

67. McGuire, K.J., McDonnell, J.J., Weiler, M., McGlynn, B.L., Kendall, C., Welker, J.M., and Seibert, J. (2005) The role of topography on catchment-scale water residence time. Water Resource Research. 41: 1-14.

68. Kohn, M. and Welker, J.M. (2005) Precipitation δ18O-temperature relations: A reexamination. Earth and Planetary Sciences Letters. 231: 87-96.

69. Dutton, A. L., Wilkinson, B.H., Welker, J.M., and Lohmann, K.C. (2005) Comparison of river water and precipitation δ18O across the 48 contiguous United States. Hydrological Processes. 19: 3551-3572.

70. Bowling, D.R., McDowell, N.G., Welker, J.M., Bond, B.J., Law, B.E., and Ehleringer, J.R. (2003) Oxygen isotope content of CO2 in nocturnal ecosystem respiration. 1. Observations in forests along a precipitation transect in Oregon, U. S. A. Global Biogeochemical Cycles. 17: 4, 1120.

71. Bowling, D.R., McDowell, N.G., Welker, J.M., Bond, B.J., Law, B.E., and Ehleringer, J.R. (2003) Oxygen isotope content of CO2 in nocturnal ecosystem respiration. 2. Short-term dynamics of foliar and soil component fluxes in an old-growth ponderosa pine forests. Global Biogeochemical Cycles. 17: 4, 1124.

72. Larson, R.M., Todd, L.C., Kelly, E.F., and Welker, J.M. (2001) Carbon isotopic analysis of bison dentition. Great Plains Research. 11: 25-64.

73. Welker, J.M. (2000) Isotopic (δ18O) characteristics of weekly precipitation collected across the United States: An initial analysis with application to water source studies. Hydrological Processes. 14: 1449-1464.

74. Harvey, F.E., and Welker, J.M. (2000) Stable isotopic composition of precipitation in the semi-arid north-central portion of the US Great Plains. Journal of Hydrology. 238: 90-109.

75. Dodd, M.B., Lauenroth, W.K., and Welker, J.M. (1998) Differential water resource use by herbaceous and woody plants in a shortgrass steppe community. Oecologia. 117: 504-512.

76. Welker, J.M., Heaton, T.H.E., Sprio, B. and Callaghan, T.V. (1995) Indirect effects of winter climate on the δ13C and the δD characteristics of annual growth segments in the long-lived, Arctic clonal plant Cassiope tetragona. Palaeoclimate Research. 15: 105-120.

77. Welker, J.M., Gordon, D.R., and Rice, K.J. (1991) Capture and allocation of nitrogen by Quercus douglasii seedlings in competition with annual and perennial grasses. Oecologia. 87: 459-466.

78. Welker, J.M., Rykiel, E.J., Briske, D.D. and Goeschl, J.D. (1985) Carbon import among vegetative tillers within two bunchgrasses: Assessment with carbon-11 labeling. Oecologia. 67: 209-212.

Plant-Animal Interactions

79. Cahoon, S., Sullivan, P.F., Post, E., and Welker, J.M. Herbivores constrain Arctic CO2 uptake and surpress C cycling responses to warming. Global Change Biology. (In Press)

80. Chimner, R., and Welker, J.M. Influence of grazing and precipitation on ecosystem carbon cycling in a Mixedgrass Prairie. Pastorial Science (In Press)

81. Chimner, R., Welker, J.M., Morgan, J., LeCain, D. and Reader, J. (2010) Experimental manipulations of winter snow cover and summer rain influence ecoystem C cycling in a Mixedgrass Prairie. Ecohydrology. 3: 284-297.

82. Ingram, L., Stahl, P., Vance, G., Schuman, G., Ganjegunte, G.K., and Welker, J.M. (2007) Grazing and drought effects on carbon, nitrogen and microbial communities in a mixedgrass prairie. Soil Science Society of America Journal. 72: 939-948.

83. Ganjegunte, G.K., Vance, G.F., Preston, C.M., Schuman, G.E., Ingram, L.I., Stahl, P.D., and Welker, J.M. (2005) Soil organic carbon composition in a northern mixedgrass prairie: Effects of grazing. Soil Science Society of America Journal. 69: 1746-1756.

84. Welker, J.M., Fahnestock, J.T., Povirk, K., Bilbrough, C. and Piper, R. (2004) Carbon and nitrogen dynamics in a long-term grazed alpine grassland. Arctic, Antarctic and Alpine Research. 36: 10-19.

85. Alstad, K.P., Welker, J.M., Williams, S. and Trilica, M.J. (1999) Carbon and water relations of Salix monticola in response to winter browsing and changes in surface water hydrology: An isotopic study using δ13C and δ18O. Oecologia. 120: 375-385.

86. Welker, J.M. and Menke, J.W. (1990) The influence of simulated browsing on tissue water relations, growth and survival of Quercus douglasii (Hook and Arn.) seedlings under slow and rapid rates of soil drought. Functional Ecology. 4: 807-817.

87. Welker, J.M., Briske, D.D., and Weaver, R.W. (1987) Nitrogen-15 partitioning within a three generation tiller sequence of the bunchgrass Schizachyrium scoparium: Response to selective defoliation. Oecologia. 24: 330-334.

Climate Change-Ecosystem and Organismic Responses

88. Ives, S., Sullivan, P.F., Dail, R., Berg, E. and Welker, J.M. Seasonal and inter-annual patterns in the magnitudes of CO2 exchange along a hydrologic gradient in the Kenai Lowlands, AK: Feedback implications of wetland drying and vegetation succession. Ecohydrology. (In Press)

89. Blumenthal, D., Chimner, R., Welker, J.M., and Morgan, J. (2008) Increased snow facilitates plant invasion in mixedgrass prairie. New Phytologist. 112: 33-39.

90. Chimner, R. A. and Welker, J.M. (2005) Soil respiration in a mixedgrass prairie: Responses to changes in snow cover and summer precipitation. Biogeochemistry. 73: 257-270.

91. Rustad, L.E., Campbell, J., Marion, G.M., Norby, R., Mitchell, M., Hartley, A., Cornelissen, J., Gurevitch, J. and GCTE-NEWS (Network of Ecosystem Warming Studies (Welker, J.M. and 15 others). (2001) A meta-analysis of the response of soil respiration, net N mineralization and aboveground plant growth to experimental warming. Oecologia. 126: 543-562.

92. Welker, J.M., Bowman, W., Williams, M., and Seastedt, T. (2001) Environmental change and future directions in alpine research. In: Bowman, W. and Seastedt, T. (Eds). Structure and function of an alpine ecosystem. Oxford University Press. 304-322.

93. Werkman, B.R., Callaghan, T.V., and Welker, J.M. (1996) Responses of bracken to increased temperature and nitrogen availability. Global Change Biology. 2: 59-66.

94. Zhang, Y. and Welker, J.M. (1996) Tibetian alpine tundra responses to simulated climate change: Biomass, nutrient and community responses. Arctic and Alpine Research. 28: 203-209.

95. Welker, J.M. and Briske, D.D. (1992) Clonal biology of the temperate caespitose graminoid Schizachyrium scoparium: A synthesis with reference to climate change. Oikos. 56: 357-365.

Carbon Cycling

96. Cable, J. Ogle, K., Welker, J.M. and 10 additional authors. (2011) The temperature response of soil respiration: A 7 desert synthesis. Biogeochemistry. 103: 71-90.

97. Hunt. E.R., Kelly, R.D., Fahnestock, J.T., Smith, W.K., Welker, J.M. and Reiners, W.A. (2004) Estimation of carbon sequestration by combining remote sensing and net ecosystem exchange data for northern mixedgrass prairie and sagebrush-steppe ecosystems. Environmental Management. 11: 201-210.

98. Smith, W.K., Kelly, R.D., Welker, J.M., Fahnestock, J.T., Reiners, W.A., and Hunt, E.R., Jr. (2002) Leaf-to-aircraft measurements of net CO2 exchange in a sagebrush steppe ecosystem. Journal of Geophysical Research – Atmospheres. 108(D3): 9.1-9.9. doi: 10.1029/2002JD002512.

99. Kelly, R.D., Hunt, E.R., Reiners, W.A., Smith, W.K., and Welker, J.M. (2002) Relationships between daytime carbon dioxide flux and absorbed photosynthetically active radiation for four different mountain/plains ecosystems. Journal of Geophysical Research – Atmospheres. 107: D14. doi:1029/2001JD001181.

100. Hunt, E.R., Jr., Fahnestock, J.T., Kelly, R.D., Welker, J.M., Reiners, W.A., and Smith, W.K. (2002) Carbon sequestration from remotely sensed NDVI and net ecosystem exchange. In: Muttiah, R. S. (Ed.), From Laboratory Spectroscopy to Remotely Sensed Spectra of Terrestrial Ecosystems. Kluwer: Dordrecht, The Netherlands. 161-174

101. Povirk, K.A., Welker, J.M. and Vance, G.W. (2001) Carbon sequestration in Arctic and alpine tundra and mountain meadow ecosystems. In: Follet, R., Kimble, J., and Lal, R. (Eds). The potential of U. S. grazing lands to sequester C and mitigate the greenhouse effect. Lewis Publishers. 189-228.

102. Zeller, K., Zimmerman, G., Hehn, T., Donev, E., Denny, D., and Welker, J.M. (2001) Analysis of in advertent microprocessor lag time on eddy covariance results. Journal of Applied Meteorology. 40: 1640-1646

Plant and Soil Water Relations

103. McGuire, K.J., McDonnell, J.J., Weiler, M., McGlynn, B.L., Kendall, C., Welker, J.M., and Seibert, J. (2005) The role of topography on catchment-scale water residence time. Water Resource Research. 41: 1-14.

104. Momen, B., Menke, J. W., Welker, J.M., Rice, K.J., and Chapin, F.S. (1994) Blue oak regeneration and seedling water relations in four sites within a California oak savanna. Int. J. Plant Sciences. 155: 744-749.

105. Rice, K.J., Gordon, D.R., Hardison, J.L. and Welker, J.M. (1993) Phenotypic variation in seedlings of a "keystone" tree species (Quercus douglasii): The interactive effects of acorn source and competitive environment. Oecologia. 96: 536-547.

106. Momen, B., Menke, J.W., and Welker, J.M. (1992) Tissue water relations of Quercus wislizenii seedlings in response to drought. Acta Oecologica. 13: 127-136.

107. Welker, J.M., McClelland, S., and Weaver, T.W. (1991) Soil water retention after natural and simulated rainfall on a temperate grassland. Theoretical and Applied Climatology. 44: 447-453.

108. Gordon, D.R., Welker, J.M., Menke, J.W., and Rice, K.J. (1989) Competition for soil water between blue oak seedlings and annual plants. Oecologia. 79: 533-541.